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Development Home Germination Early growth and tillering Stem elongation Wheat apex development Flowering and fertilisation Pollen release Grain growth 1 to 4 days Grain growth 4 to 10 days Cell layers inside a grain Grain filling 11 to 16 days Grain filling 17 to 21 days Whole grain photographs Endosperm micrographs Embryo micrographs
Grain filling 21 to 30 days Development of the embryo Dry down 30 to 40 days How we grew the plants Photography and microscopy
DAYS = days after flowering
Development Home Germination Early growth and tillering Stem elongation Wheat apex development Flowering and fertilisation Pollen release Grain growth 1 to 4 days Grain growth 4 to 10 days Cell layers inside a grain Grain filling 11 to 16 days Grain filling 17 to 21 days Whole grain photographs Endosperm micrographs Embryo micrographs
Grain filling 21 to 30 days Development of the embryo Dry down 30 to 40 days How we grew the plants Photography and microscopy
DAYS = days after flowering
Grain filling 17 to 21 days
Whole grain photographs
 | Whole grain at the soft dough stage showing the ventral groove |
 | Dorsal view of the whole grain at the soft dough stage. The grain can still be squashed with a thumbnail but the contents are much firmer than during the Milk stages. It is half way through the grain filling stages. |
 | An individual spikelet 24 days after pollination. The grain has grown and is changing colour. The glumes, lemmas and palea, which protect the grain are drying off and losing their green colour. floret 1 has been cut away and its component parts are separated. |
 | The outer layers of this grain have been dissected away to show; 1)the maternal pericarp is changing colour from green to yellow 2)the embryo is about half its final size. The embryo unit is about 2.0mm long but much of this is the plate of the scutellum which is attached to the tiny embryo and provides it with nutrients from the endosperm. You can see this better in the embryo sections at 20 days after flowering. |
 | Longitudinal section of the fresh grain through the ventral groove. The embryo occupies its final position near the base of the grain against the dorsal surface. The embryo is about 2.0mm long and had primordial leaves within the coleoptile at the shoot pole. |
 | Transverse section of the fresh grain at maximum fresh weight. The endosperm has filled out and the maternal epidermis is a closer fit on the dorsal side. The endosperm still contains some water at this stage. |